A couple of things about the Tkrai...

When writing about them you should attend to how their biological differences would effect their culture. This is a better approach than copying Earth cultures would be, because whereas copying earth cultures is easy what you will end up with will be pretty nooby.

We've established that, in their native environment when they are not forced to live under human conditions, the Tkrai are climbers. Their physiology is gonna most likely be some sort of cross between monkeys, bats, and flying squirrels, with an emphasis on the bat-look. They have proto-wings with which they most likely cannot fly, but they could glide from tree to tree or rock to rock. This is also going to change the way they fight, since they wouldn't use their limbs in the same exact ways that humans do. I don't think spears will necessarily be that popular. I've introduced the use of obsidian blades that are strapped to the wrists or ankles as something that exists in the south. Whether or not this also exists in the north has not been decided. I also figure that their warfare, even in the context of advanced civilizations, won't look anything like human warfare. Tkrai legions lining up and fighting in formation seems unwieldy. We're most likely looking at something that is more three dimensional and organized in a way that isn't necessarily easy to compare to human fighting. A well orchestrated guerrilla-skirmish type warfare that involved every surface available. 

Their psychology is going to be different too. This is a race that lives on a planet so aggressively difficult to survive that their are practically at war with the environment. After all, we've established that humanity is having a hard time pushing back the wilderness, and has been practically incapable of doing so past the safe zone. They are going to be hardy. 

Another thing to remember is that Googer and Hugs have established this world as being porous and stringed with caves. There is going to be a world underground that we haven't explored yet, and I suspect the Tkrai will be a major part of that.